My research interests include developmental and cytogenetic studies in amphibians, especially salamanders, including evolutionary cytogenetics and genomics, genome size, limb and organ regeneration, and amphibian deformities. Recent work has involved international collaborations with colleagues in Japan, Slovenia, England, Russia, and Canada. Current research includes cytogenetics and limb developmental evolution in Asian hynobiid salamanders, genetics, cytogenetics, and developmental evolution of cave adaptations in the European blind cave salamander (Proteus anguinus), and genome size evolution and its phenotypic correlates in salamanders, especially Neotropical plethodontids.